Why do the bees fail to completely cap or open some pupae?
(http://www.wjclemens.com/cordovan-honeybee/general_hive_photos/Feral_Survivor_brood_NucD_marked.jpg)
I think the pupa have died and they know it , they sense no movement and the dead pupa have a smell if you will mark those uncapped cells, for a short time you will find them empty.
The whole concept of hygenic bees is that they uncap pupae that are not perfect in some way. If the pupae is infested with mites or infected with some brood disease they uncap and then remove them.
But they have isolated several genes that are tied to that behavior. Some have to do with uncapping and some have to do with removing. If a lot of it stays open like that, maybe you have one and not the other.
Thanks Michael,
I'm going to take a few follow-up photo's and see how the bees have dealt with these.
Joseph,
an interesting read
http://www.beekeeping.com/articles/us/hygiene_queen.htm
Dave